Table 1

In vivo murine mammary morphogens

Morphogen
Branching*
Mode of application
Reference

Extracellular factors



     Estrogen

Administered
[110]
     Estrogen

Slow-release pellet
[111]
     Anti-estrogen

Slow-release pellet
[112]
     Progesterone

Slow-release pellet
[113]
     Parathyroid-hormone-related peptide

Homozygous deletion
[114]
     Parathyroid-hormone-related peptide

Transgenic overexpression
[115]
     Parathyroid hormone

Transgenic overexpression
[115]
     Corticosterone

Administered
[116]
     Epidermal growth factor (EGF)

Slow-release pellet
[117]
     TGF-β1

Slow-release pellet
[118-119]
     TGF-β2

Slow-release pellet
[118]
     TGF-β3

Slow-release pellet
[118]
     Insulin-like growth factor 1

Homozygous deletion
[120]
     Growth hormone

Slow-release pellet
[117]
     Hepatocyte growth factor

Transgenic overexpression
[121]
     Relaxin

Systemically administered
[122]
     Amphiregulin

Overexpressing transplants
[123]
     Amphiregulin

Slow-release pellet
[123]
     Hereguln-α

Slow-release pellet
[124]
     Heregulin-β

Slow-release pellet
[124]
     Wnt-4

Homozygous deletion
[125]
     Wnt-4

Overexpressing transplants
[126]
     C-neu

Transgenic overexpression
[127]
     Colony-stimulating factor

Homozygous deletion
[128]
     Eotaxin

Homozygous deletion
[128]
     MMP-3 (stromelysin-1)

Transgenic overexpression
[69-70]
     TIMP-1

Transgenic antisense TIMP-1
[12]
     TIMP-1

Slow-release pellet
[12]
Receptors



     EGF receptor (EGFR)

Homozygous deletion
[129]
     Epidermal-growth-factor receptor

Transgenic/dominant negative
[130]
     Estrogen receptor α

Homozygous deletion
[131]
     Progesterone receptor A (PRA)

Transgenic overexpression
[132]
     Progresterone receptor A/B

Homozygous deletion
[133]
     Prolactin receptor

Homozygous deletion
[134]
     TGF-β2 receptor

Transgenic/dominant negative
[135]
Intracellular factors



     CCAAT/enhancer binding protein β

Homozygous deletion
[136-137]
     Gelsolin

Homozygous deletion
[138]
     Hox6C

Homozygous deletion
[139]

* ↑, augmentation of branching; ↓, inhibition of branching. MMP, matrix metalloproteinase; TGF, transforming growth factor; TIMP, tissue inhibitor of metalloproteinase.

Fata et al. Breast Cancer Res 2004 6:1   doi:10.1186/bcr634